Job openings

    • 31 Aug 2022
    • Basel, ETH Zurich

    This is an interdisciplinary Post-Doc position funded by a 3-year grant from the Personalised Health and Related Technologies (PHRT) program. You will work in the group of Prof Iber (ETH, computational biology), and collaborate with Prof Ulbrich (ETH, animal physiology), and Prof De Geyter (University Hospital Basel, reproductive medicine). 

    Project background

    The goal of the project is to develop computational personalized medicine approaches to improve the success rate of in vitro fertilization (IVF) using human and farm animal data. Given our detailed understanding of ovarian follicle maturation, personalized medicine approaches can go beyond statistical correlation and can be based on a mechanistic understanding of the hormonal control of ovarian follicle maturation.

     Job description

    You will focus on the development of predictive mathematical models that integrate literature data and newly generated data from farm animals and patients (3D images, hormone levels, gene expression time series) into a consistent framework that will allow us to provide data- and model-driven guidance regarding the best IVF treatment schedule. In collaboration with a PHRT-funded PhD student, the mathematical model will be carefully tested in clinical settings and then integrated into software for clinical use.

    Your profile

    • You are experienced in building mechanistic, predictive mathematical models based on biological data.
    • It is a plus if you have previous experience with image processing, finite element methods, and software engineering, as well as expertise regarding the female endocrine and reproductive system.
    • You are self-motivated and enthusiastic about leveraging mechanistic modelling for personalised medicine, advancing FemTech, and contributing to the establishment of a strong personalised medicine/personalised health research community in Switzerland, for the eventual benefit of patients and citizens.

    We offer

    • an internationally highly competitive salary and further benefits
    • excellent working conditions, with the possibility to work partly in the home office
    • a friendly and collaborative working environment
    • the opportunity to translate science to the clinic
    • support for career development after the PHRT-funded Post-Doc via suitable academic fellowships such as SNF Ambizione, or an ETH Pioneer Fellowship to commercialise the results

    The preferred starting date is October 2022; the exact starting date can be negotiated. 


    Please apply via our online application portal: 

     Further information about the group can be found on our website: Questions regarding the position should be directed to Prof Iber by e-mail: (no applications).

    • 1 Sep 2022
    • Autonomous University of Barcelona (UAB), Spain

     1.Basic job and project description:
    The position is the Autonomous University of Barcelona (UAB), Spain

    -The main project questions are:
    How gene networks evolved to lead to the evolution of complex and robust phenotypes?

    Can be uncover some principles on how gene networks wired over evolution to reach such complexity?

    How these principles affect the tempo and mode of evolution.

    Any other question of the applicants interest that are related to the previous questions.

    We will use computational models of gene networks during development and computational models of evolution by natural selection to study such questions for the case of organismic complexity (e.g. anatomy). The models we develop are genotype-phenotype maps, relating genetic changes to phenotypic variation. These models are based on realistic gene network models of pattern formation and morphogenesis that lead to realistic 3D multicellular (animal) phenotypes. So actual embryonic and adult morphologies arise from our models, either of specific organs (e.g. teeth, wings, limbs) or generic embryo-like morphologies.

    The position is in Salazar-Ciudad’s group and it entails obtaining a PhD degree.

    2. Background of the project:
    There is no consensus definition of complexity, yet it is evident that organisms are complex and explaining such complexity is one of the most fundamental questions of biology. Morphological complexity has not increased in the evolution of all lineages and, in general, it is unclear whether there is a general trend of increasing complexity in evolution. Yet, one may ask about the mechanisms by which such complexity has increased in the lineages where it has increased. How complexity increases during evolution is necessarily related to development: any evolutionary change in morphology is first a change in the development that produces such morphology.

    It has been argued that, in spite of the remarkable morphological complexity of organisms, their development is achieved through a limited number of cell behaviors and types of cell interactions. These cell behaviors would be cell division, cell adhesion, cell death, cell growth, cell contraction, extracellular signal and matrix secretion, extracellular signal reception and cell differentiation. In addition to cell behaviors, development involves interactions between cells, either mechanical or through extracellular signalling.

    The questions we want to approach in this study are: how should these interactions and cell behaviors be coordinated to produce complex and robust morphologies in evolution? The question is, then, whether there are some logical requirements that developmental mechanisms should fulfill in order to lead to complex robust morphologies. Are there, for example, some requirements at the level of gene network topology or at the level of cell behaviors and their coordination during development? How do these requirements arise in evolution?

    If, as suggested above, pattern transformations in development involve a limited set of cell behaviors and cell interactions, then any mathematical model implementing those and intracellular gene networks should be able to reproduce, to a large extent, the range of pattern transformations possible in animal development. In this project we will use one such model, EmbryoMaker (Marin-Riera et al, 2015)⁠, to simulate a large number of possible developmental mechanisms and try to discover what, if anything, do the mechanisms leading to robust complex morphologies have in common.

    3. Job description
    The main tasks of the student include using existing models of embryonic development (e.g. EmbryoMaker) in order to simulate the development of complex morphologies. These models will be combined with models of evolution, in a population context with mutation, genetic drift and natural selection on morphology (see for example Salazar-Ciudad and Marin-Riera, 2013). The gene networks found to lead to the development of complex and robust phenotypes in evolution would be analyzed to extract general regularities, if any, these gene networks need to fulfill. The main tasks, thus, consist in simulation, theorizing, data analysis, coding, literature searching, writing and presenting results in conferences. We seek candidates highly motivated for theoretical work and data analysis with a broad understanding of the evolutionary theory and/or developmental biology and/or modeling.

    4. Requirements:
    - Candidates should have a University Degree and a Master’s Degree in biology, or similar, within the European Higher Education System (minimum 300 ECTS) or equivalent by September 2021.

    -Scientific programming skills or a willingness to acquire them is required.

    -The most important requirement is a strong interest and motivation on science and evolution. A capacity for creative and critical thinking is also required.

    5. Salary and conditions:
    -The salary would be the standard one graduate students in Spain. Full-time employment

    6. The application must include:
    -Application letter including a statement of interests and motivation.

    -CV (including publications, the contribution of the applicant in each publication, degrees obtained, subjects included in degree and grades, average grade)

    Foreign applicants, especially non-EU applicants, should attach an explanation of their University's grading system. All documents should be in English, no official translation is required in the initial application but may be requested afterwards. Applications should be sent to Isaac Salazar-Ciudad by email:

    7. Starting:
    The start date is negotiable after September of  2022.

    8. Application deadline:
    1st of September 2022

    9. Work environment:
    Salazar-Ciudad group is between the Genetics deparment in the UAB ( and the mathematics research center, CRM:

    Both the CRM and Genetics department are very international and, thus, not being able to speak Catalan or Spanish is not a problem. Nearly half of the PIs in the CRM are foreigners. The everyday working language is English, and most administrative tasks, training and seminars arranged by the department and CRM are conducted in English.

    The UAB is one of the largest and best Universities in Spain.

    10. References:
    Salazar-Ciudad I, Marín-Riera M.Adaptive dynamics under development-based genotype-phenotype maps. Nature. 2013 May 16;497(7449):361-4.

    Hagolani PF, Zimm R, Marin-Riera M, Salazar-Ciudad I. Cell signaling stabilizes morphogenesis against noise. Development. 2019 Oct 18;146(20):dev179309. doi: 10.1242/dev.179309. PMID: 31628213

    Hagolani PF, Zimm R, Vroomans R, Salazar-Ciudad I. On the evolution and development of morphological complexity: A view from gene regulatory networks. PLoS Comput Biol. 2021 Feb 24;17(2):e1008570.

    • 2 Sep 2022
    • University of Udine, Italy

    Open call for PhD position on “Computational methods for analyzing the dynamics of certain diseases” in the PhD programme in Mathematical and Physical Sciences at the University of Udine under the funding of the National Recovery and Resilience Plan (PNRR).

    The successful candidate will join the research group at CDLab ). 

    You can find more details at

    (section Mathematical and Physical Sciences​)

    • 8 Sep 2022
    • Krakow, Poland

    I am looking for two students interested in pursuing their PhD in Biophysics focused on understanding the biophysical limiting factors of patterning precision in developing tissues. The PhD studies will take place at the Faculty of Physics, Astronomy and Applied Computer Science of Jagiellonian University, Kraków, Poland. The positions are fully funded by the National Science Center Poland (NCN).

    The project goal is to understand to what extent growth, cellular dynamics, biomechanical feedbacks and global mechanical constraints limit patterning precision in the growing tissue. The successful candidate is expected to actively take part in the project by proposing and analyzing models in the framework of theoretical and computational biophysics. The proposed models will be informed with high resolution experimental data provided by external collaborators.

    The candidate is expected to have:

           MSc (or equivalent) in biophysics, physics, mathematics, computer science or similar, allowing application to the Doctoral School,

           excellent academic track record,

           interest in the interdisciplinary aspect of the project,

           experience with numerical solvers (C++, Python, Mathematica),

           proficiency in written and spoken English.

    We offer:

    • interdisciplinary research on interface of physics and biology,
    • research projects jointly with experimental labs (Austria, Poland),
    • opportunities of research visits in collaborating labs and participation in international conferences.

    The exact requirements, dates and details on application procedure are available at the webpage of Doctoral School of Exact and Natural Sciences

    I encourage interested candidates to contact me in advance at and provide CV, list of publications, and at least one recommendation letter. Further information on group activity is available at

    The application deadline is September 8, 2022.

     Marcin Zagórski

    • 9 Sep 2022
    • INRIA Saclay - Île-de-France

    Recent biological imaging have shown that membrane receptors involved in intracellular signaling pathways are capable of inducing cascades of biochemical reactions from extremely dynamic pools of intracellular compartments. The subcellular trafficking of receptors generates spatio-temporal heterogeneous cellular compartments with a critical role on physiological functions, and profound consequences on the search for new therapeutic strategies.

    The candidate will develop new modeling formalism, combining biochemical reaction networks and coagulation-fragmentation dynamics to fully represent the diversity and complexity of signaling pathways. By comparing the models with data from confocal microscopy imaging with super spatio-temporal resolution, the candidate will reveal how the spatio-temporal heterogeneity of intracellular compartments conditions the response of cells to extracellular signals.

    The candidate will design efficient numerical schemes for partial differential equation (PDE) of coagulation type to explore the role of endocytosis, recycling and coagulation-fragmentation dynamics in receptor trafficking, as well the importance of highly dynamics phase transition and coarsening phenomena in downstream second messenger activation.

    Using ideas from chemical reaction network theory, the candidate will analyse the long-time behavior of the developed models, according to the underlying reaction network topology and coagulation-fragmentation dynamics, and its consequence for cellular response.

    Please contact Romain Yvinec ( for more information and before applying.

    Detailed annoucement and application procedure:

    • 15 Sep 2022
    • Spain

    The @Mathematical and Theoretical Biology Group at BCAM is hiring promising researchers!!! 2 positions to be filled.

    Please see information below:

    - 1+ 1 years Postdoctoral Researcher on "Modeling spatial dynamics of disease spreading". Within the BMTF project, the research project will be focused on the development of spatio-temporal models variations on infectious diseases transmission and control and its impact on the Basque surveillance system; Modeling different social contact structures to understand disease spreading; Modeling the role of social behavioral changes, lock downs and other control measures affecting the dynamics of seasonal respiratory disease.

    Applicants must have a relevant curriculum in the scientific area of biomathematics applied to infectious disease, stochastic processes and network theory. We look for applicants with PhD degree in applied mathematics, mathematical biology, physics or related disciplines on quantitative life sciences field.

    Apply here:

    Deadline: September 15th 2022, 14:00 CET (UTC+1) .

    - 1 year Research Technician position to work on "Modeling infectious disease dynamics"

    We seek promising young researchers, eventually interested to develop a PhD project on infectious disease dynamics.

    Note that a PhD position is to be financed by a national or international call. Candidates must apply for the PhD grant individually, with the support of the host institution and supervisor.

    The preferred candidate will have: 
    • Strong background in statistics
    • Information Technology Background
    • Knowledge of bio-statistics and experience in advanced programming are essential.

    Applicants must have their Bachelor’s or Master degree preferable in computer science, mathematics or quantitative life sciences field.

    Apply here:

    Deadline: September 15th 2022, 14:00 CET (UTC+1) .

Previous openings

5 Aug 2022 3-year Postdoc position in Theoretical Evolutionary Ecology
31 Jul 2022 PhD position in Theoretical Biology
13 Jul 2022 PhD scholarship on mathematical modelling of brain development and evolution
10 Jul 2022 Postdoctoral Research Fellow
8 Jul 2022 Research scientist positions in mathematical biology at RIKEN
5 Jun 2022 Postdoc Antimicrobial resistance modeling for global health interventions
25 May 2022 Research Software Engineers (x2)
4 May 2022 PhD position in the Theoretical Biology group at Bielefeld University
29 Apr 2022 PhD scholarship: Mathematical modeling in infection and immunity
25 Apr 2022 Two 3-Year Post-Doc positions at the interface of developmental biology and mathematical modelling
20 Apr 2022 PhD project in Systems Biology
20 Apr 2022 Postdoctoral Researcher
20 Apr 2022 Professorship (W2 or W3) on Modelling in the life sciences
17 Apr 2022 Lecturer in Mathematical Biology
15 Apr 2022 PhD position: Ecosystem-based and evolutionary approach to the exploitation of species
15 Apr 2022 PhD position: Mathematical theory of tumour evolutionary modes
15 Apr 2022 PhD Position: Multi-scale mathematical modeling of oogenesis in fish
27 Mar 2022 Postdoctoral and Senior Postdoctoral Research Fellow Positions in Cardiac Electrophysiology Modelling and Inference
2 Mar 2022 PhD position: Mathematical modelling and data driven analysis of cooperation in stochastic games
28 Feb 2022 IBS Biomedical Math Senior Researcher & Postdoc Fellow
15 Feb 2022 Two Postdoc Positions in Mathematical Modeling of Microbial Evolution
8 Feb 2022 PhD positions in Evolutionary Biology
31 Jan 2022 5 senior and 4 junior postdoctoral at the Centre de Recerca Matemàtica (CRM) in Barcelona
27 Jan 2022 6 PhD fellowships - doctoral fellowship programme INPhINIT “la Caixa”
21 Jan 2022 PhD position in Evaluating the effectiveness and cost-effectiveness of one-health vaccination strategies
19 Jan 2022 Two PhD positions in Theoretical Biology
11 Jan 2022 Tenure-track Faculty Position in Numerical Analysis
15 Dec 2021 Assistant Professor of Mathematical Health and Disease Modelling
6 Dec 2021 Postdoctoral Research Associate in the Topological Data Analysis of Lung Cancer
1 Dec 2021 Postdoc position modeling and analysis of PDEs
30 Nov 2021 Research Fellow (3-year postdoc) in Mathematical modelling of the Bunyaviruses intra-cellular life cycle
27 Oct 2021 Postdoc position: Modelling Individualisation in Evolution and Economics
11 Oct 2021 Postdoc position at the Schumacher group in Edinburgh
10 Oct 2021 PhD position in data-driven mathematical modelling for microbiology
7 Oct 2021 Postdoctoral Fellow Position in Multiscale Modeling of Embryonic Development
3 Oct 2021 2 posts as Lecturer in Applied Mathematics
30 Sep 2021 Postdoctoral Researcher at the Mathematical and Theoretical Biology Group, BCAM
8 Aug 2021 Senior Researcher positions
2 Aug 2021 Research Technician in Biomathematics at the Mathematical and Theoretical Biology Group, BCAM.
30 Jul 2021 PhD studentship: An eco-evolutionary framework for studying resilience and tipping points in ecosystems under stress
30 Jun 2021 Postdoc position on biophysical modelling
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20 Jun 2021 PhD position in mathematical systems biology
31 May 2021 Funded positions at Trento and Verona
17 May 2021 PhD Position in Mathematical Biology
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13 May 2021 PhD positions in Pure and Applied Mathematics.
10 May 2021 Research Fellow in Modelling and Analysis of Cancer Invasion
30 Apr 2021 Cluster Hire in Quantitative and Computational Developmental Biology
30 Apr 2021 RA or Postdoc on modeling COVID-19 for applied public health at INSERM
28 Apr 2021 Postdoctoral Research Associates, University of Oxford
16 Mar 2021 Research fellowship positions at IBS Biomedical Mathematics Group, Daejeon, Korea
13 Mar 2021 Assistant Professor at Heriot-Watt University, Edinburgh
11 Mar 2021 Post-Doc in Evolutionary Theory and Disease at U Maryland
8 Mar 2021 Postdoctoral position at the Mathematical and Theoretical Biology Group at BCAM
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24 Feb 2021 PhD studentship in mathematical modelling of cancer evolution at City, University of London
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14 Feb 2021 PhD Fellowship at the Delft University of Technology
10 Feb 2021 BCAM - Group Leader position in Mathematical Modelling applied to Health
29 Jan 2021 Research fellow positions in the Mathematical and Theoretical Biology group at BCAM
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27 Feb 2020 PhD position at the Wilhelmina Children's Hospital in Utrecht.
17 Feb 2020 PhD position in Applied Mathematics within the joint graduate programme of Swinburne University of Technology in Melbourne, Australia and Politecnico di Torino, Italy.
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8 Jan 2020 Three Fully Funded PhD positions in Quantitative Developmental Biologyand Mathematical Modeling at Leiden University, The Netherlands
19 Dec 2019 PhD position at the University of Bath
11 Dec 2019 PhD position at Bielefeld University
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30 Oct 2019 PhD position in Mathematical Biology at the Max Planck Institute for Evolutionary Biology.
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1 Jul 2019 1 Assistant Professor and 2 Junior Faculty Positions at Department of Mathematics, University of California, Riverside (USA)

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